The common loon or great northern diver ( Gavia immer) is a large member of the loon, or diver, family of birds. Reproduction adults have a plumage that includes a broad black head and neck with a greenish, purplish, or bluish sheen, blackish or blackish-grey upperparts, and pure white underparts except some black on the and vent. Non-breeding adults are brownish with a dark neck and head marked with dark grey-brown. Their upperparts are dark brownish-grey with an unclear pattern of squares on the shoulders, and the underparts, lower face, chin, and throat are whitish. The sexes look alike, though males are significantly heavier than females. During the breeding season, loons live on lakes and other waterways in Canada, the northern United States (including Alaska), and southern parts of Greenland and Iceland. Small numbers breed on Svalbard and sporadically elsewhere in Arctic Eurasia. Common loons winter on both coasts of the US as far south as Mexico, and on the Atlantic coast of Europe.
Common loons eat a variety of animal prey including fish, , insect larvae, Mollusca, and occasionally aquatic plant life. They swallow most of their prey underwater, where it is caught, but some larger items are first brought to the surface. Loons are monogamous; that is, a single female and male often together defend a territory and may breed together for a decade or more. Both members of a pair build a large nest out of dead Hymenachne and other plants formed into a mound along the vegetated shores of lakes. A single brood is raised each year from a clutch of one or two olive-brown oval Bird egg with dark brown spots which are Egg incubation for about 28 days by both parents. Fed by both parents, the chicks fledge in 70 to 77 days. The chicks are capable of diving underwater when just a few days old, and they fly to their wintering areas before ice forms in the fall.
The common loon is assessed as a species of least concern on the IUCN Red List of Endangered Species. It is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds applies. The United States Forest Service has designated the common loon a species of special status because of threats from habitat loss and toxic metal poisoning in its US range.
The common loon is the provincial bird of Ontario, and it appears on Canadian currency, including the Loonie coin and a previous series of $20 bills. In 1961, it was designated the state bird of Minnesota, and appears on the Minnesota State Quarter and the state Seal of Minnesota.
Danish zoologist and mineralogist Morten Thrane Brünnich first described the common loon in 1764, as Colymbus immer in his Ornithologia Borealis. The now-defunct genus Colymbus contained as well as loons, and remained in use until the International Commission on Zoological Nomenclature attempted to clarify the nomenclature in 1956 by declaring Colymbus a suppressed name unfit for further use and establishing Gavia, created by Johann Reinhold Forster in 1788, as the valid genus name for the loons.
The current genus name Gavia was the Latin term for an unidentified seabird and the specific immer is derived from a Norwegian name for the bird, similar to the modern Icelandic word "himbrimi". The word may be related to Swedish immer and emmer: the grey or blackened ashes of a fire (referring to the loon's dark plumage); or to Latin immergo, to immerse, and immersus, submerged.
The European name "diver" comes from the bird's practice of catching fish by diving. The North American name "loon" was first recorded in this sense in New Englands Prospect (1634) by William Wood (1580–1639); "The Loone is an ill shap'd thing like a Cormorant". It may be derived from Old Norse lómr, as are modern Swedish and Danish lom, in each case referring to the distinctive call.
A number of fossil loon species are known from the Pliocene, and specimens from the Pleistocene of California and Florida appear to represent a paleosubspecies of the common loon.
Adult non-breeding plumage is brownish with a dark neck and head marked with dark grey-brown. The eyes are surrounded with white, and the eyelids are pale. The bill is mostly pale grey, with a dark culmen and tip, but in early spring the tip may turn whitish. The underparts, lower face, chin, and throat are also whitish. The foreneck is whitish, usually forming wedge-shaped notch in dark neck-sides, and may sometimes reveal a shadowy trace of the neck ring or a pale collar. It has dark brownish grey upperparts with an unclear pattern of squares on the shoulders and some wing coverts spotted with white, which are usually concealed while swimming. The male and the female have similar appearances, although they exhibit sexual dimorphism in their physical dimensions with the male larger and significantly heavier than the female.
The heavy dagger-like bill is evenly tapered and greyish, sometimes having a black tip. The bill colour and angle distinguish this species from the yellow-billed loon. The neck is short and thick. The common loon swims very low in the water, with sometimes only its head held above and horizontal to the surface. It must run across the water surface to get in flight. During flight, its head is slightly lower than its body, with its feet trailing behind. It has a skeletal structure made up of a number of solid bones (this is usual for the Gaviiformes and penguins but unlike most flying birds which have bird skeleton with extensive pneumatization, hollow and filled with air to make the skeleton lighter), which adds weight but helps in diving. A juvenile often has a dark, brownish-grey nape that may look darker than the pale-edged black feathers. It has a dark grey to black head, neck, and upperparts, with white throat, cheeks, and underparts. During the first winter, the bill shape of the young may not be as fully developed as that of the adult, and during the second winter, it much resembles the breeding adult, but with wing coverts lacking white spots.
The common loon is distinguished from the black-throated loon ( G. arctica) and the red-throated loon ( G. stellata) mainly by its larger size. It usually has a steeper forehead and a bulging forecrown, somewhat similar to the black-throated loon. Its bill is heavier and the back is paler than its hindneck. It is more difficult to separate from the yellow-billed loon, but its breeding plumage has more white markings on the neck and the squares on its shoulders are usually smaller; the non-breeding plumage has darker neck-sides contrasting more sharply with pale areas and bill colour.
They usually winter along coasts and on inland lakes, bays, inlets, and streams, with birds migrating to the nearest body of water that will not freeze over in the winter: western Canadian loons go to the Pacific Ocean, Great Lakes loons to the Gulf of Mexico region, eastern Canadian loons to the Atlantic, and some loons to large inland lakes and . They appear in most of the inland waters of the United States. The South Carolina coast, the Gulf coast adjacent to the Florida panhandle, and the Atlantic seaboard from Massachusetts to Maine have some of the highest concentrations of common loons. Occasional vagrants are recorded inland in Mexico, in San Luis Potosí and Coahuila, as well as in Chiapas and Oaxaca in the south. They are accidental in northern Japan and the Commander Islands in northwestern Pacific, and Cuba in the West Indies.
Common loons sometimes form social gatherings of several adults in late summer. Studies in North America suggest that these events are attended largely by non-breeding individuals and relate to territory assessment and social familiarity rather than foraging or migration.
The common loon uses its powerful Hindlimb to propel its body underwater at high speed to catch its prey, which it then swallows head-first. If the fish attempts to evade the common loon, the bird chases it down with excellent underwater manoeuvrability due to its strong legs. Most prey are swallowed underwater, where they are caught, but some larger prey are first brought to the surface. It is a visual predator, so it is essential to hunting success that the water is clear. It normally dives , but has been recorded to dive up to . The average diving time is 42 seconds, but the maximum duration spent underwater is about .
Pairs do not remain together during winter; in addition, males usually precede females by a few days to a few weeks during spring migration, settling on their lake once a portion of it becomes ice-free. Copulation takes place ashore, often on the nest site, repeated daily until the eggs are laid. The preceding courtship is very simple, with mutual bill-dipping and dives. The displays towards intruders, such as bow-jumping (an alternation of fencing and bill-dipping postures and rushing (running "along the surface with its wings either folded or half-extended and flapping at about the same speed as when taking off") are often misinterpreted as courtship.
Nesting typically begins in early May. Significantly more nesting sites are found on islands than on mainland shoreline. Breeding pairs patrol their territories routinely, even at night, defending the territory both physically and vocally. Pairs that nested together the preceding year typically reuse the nest site from the previous year, if they hatched chicks successfully there. In contrast, pairs that lost their eggs to a predator usually shift the nest to a new location. This logical behaviour pattern appears to depend upon the male, because breeding pairs consisting of last year's male and a female not present during the preceding year continue to exhibit the behaviour; pairs composed of last year's female and a new male tend to select a new nest site, regardless of the success or failure of the previous year's attempt. Despite the lead role of males in nest site selection, both sexes contribute substantially to nest construction. The nest is about wide and is constructed out of dead Hymenachne and other indigenous plants, and formed into a mound along the vegetated coasts of lakes greater than . After a week of construction in late spring, one parent climbs on top to mould the interior of the nest to the shape of its body. Based on a number of studies, nesting success averages about 40%, and most newly hatched young survive due to parental care.
Eggs from first clutches are typically laid in May or early June, the timing depending largely upon the date that lakes become ice-free and inhabitable. A clutch consists of two (occasionally one) olive-brown oval with dark brown spots. Incubation is carried out jointly by male and female and lasts about 28 days. Loons often place nests along steep lake shorelines where adults can quickly dive underwater when approached by predators. The eggs are about long and wide and the two eggs are laid with an interval of one to three days between them, and hatch .
Newly hatched chicks are dark chocolate brown in colour and have a white belly. Within hours of hatching, the young begin to leave the nest with the parents, swimming close by and sometimes riding on one parent's back. Parents and chicks initially stay in shallow, isolated bays where the parents are able to defend the chicks better from intruding loons and eagles, which are their main predators. Male parents defend broods consisting of two chicks more vigorously than singleton chicks, chiefly with the territorial yodel call. The chicks are capable of making shallow dives from their first day but make deeper dives as they grow. Fledge takes 70 to 77 days. Usually, only one brood is raised. Both parents feed the chicks live prey from hatching to fledging. As they grow, chicks are able to catch an increasingly large proportion of their diet by themselves; they can feed and fend for themselves after about two months, although many juveniles continue to beg from adults well beyond this age. The parent birds capture small fish and hold them crosswise in their bill, call and approach the chicks with their head lowered so that the chicks can grasp them. If food is scarce, the larger chick may peck its small sibling incessantly; on small lakes with limited food, only one chick often survives. Juveniles leave the breeding ground before ice formation in the fall, weeks after their parents. A pair of loons raising two chicks have been estimated to feed on of fish during the five and a half months that they spend in their breeding territory.
The tremolo call—sometimes called the "laughing" call—is characterized by its short, wavering quality. It often uses this call to signal distress or alarm caused by territorial disputes or perceived threats. It emits a tremulous series of up to 10 rather high notes (hu)-heheheheheheha. It also uses the tremolo to communicate its presence to other loons when they arrive at a lake, often when they are flying overhead. It is the only vocalization used in flight. The tremolo call has varying three levels of intensities that correlate with a loon's level of distress, and the types are differentiated by increasingly higher pitch frequencies added to the call.
The yodel is a long and complex call made only by the male. It is used in the establishment of territorial boundaries and in territorial confrontations, and the length of the call corresponds with the loon's level of aggression. The dominant frequencies in the yodel indicate the body mass and thereby the health of males. A male that occupies a new territory appears to alter its yodel to be clearly distinguishable from the call of the previous territory owner.
A loon's wail is a long call consisting of up to three notes, and is often compared to a wolf's Howling. It uses this call to communicate its location to other loons. The call is given back and forth between breeding pairs or an adult and its chick, either to maintain contact or in an attempt to move closer together after being separated. It is a loud aaoo, weee-wea weee-wea weee-wea, or ooo-aaah-éééé.
The hoot is a short, soft call and is another form of contact call. It is a more intimate call than the wail and is used exclusively between small family groups or flocks. The common loon hoots to let other family or flock members know where it is. This call is often heard when the adult loon is summoning its chicks to feed.
Eggs are taken by a number of mammals, including American mink, striped skunk, otters, foxes and , with the latter being responsible for nearly 40% of all nest failures. Birds such as , and will eat unattended eggs. Because their nests are at the water's edge, common loon eggs are especially vulnerable if the adult is absent.
Chicks may be killed by common snapping turtles, large gulls, bald eagles and large fish such as northern pike and largemouth bass. The eagle in particular is a significant predator of chicks.
Internal parasites of the common loon include many species of helminths, including trematoda, cestoda, and acanthocephala. High levels of worms may result from feeding changes due to low availability of fish, and can lead to illness and death. Protozoal infections including one caused by Eimeria gaviae and avian malaria have been recorded in this loon. The black fly Simulium is closely associated with the common loon to which it is attracted to chemicals in the uropygial gland secretions as well as by visual and tactile cues. This fly is detrimental to loons, their preferred hosts, transmitting blood-borne parasites and viruses, and causing nest abandonment when numbers are high. External parasites include bird louse, although these are not found on the bird's head.
Botulism, acquired by eating infected fish, can lead to paralysis and drowning. Aspergillosis is another cause of emaciation and death. Outbreaks sometimes lead to thousands of deaths.
The common loon is listed under Appendix II of the Convention on the Conservation of Migratory Species, and in Article I under the European Union (EU) Birds Directive. It is one of the species to which the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA) is applied. In Europe it appears in 20 Important Bird Areas (IBAs), including Ireland, Svalbard, mainland Norway, Iceland, Spain, and the United Kingdom. It is also a listed species in 83 Special Protection Areas in the EU Natura 2000 network. The USDA National Forest Service has designated the common loon a species of special status, and in the upper Great Lake regions of the Huron-Manistee, Ottawa, and Hiawatha national forests as a regional forester sensitive species.
Although the common loon is able to decrease their methylmercury levels by molting and laying eggs, continued consumption of fish with raised methylmercury levels prevents these mechanisms from effectively lowering methylmercury levels. Mercury concentrations have been shown to vary by the sex and age of common loons. Male common loons were found to contain the highest blood mercury concentration likely due to the fact that they tend to consume bigger fish with higher mercury concentrations. Females contained the second highest blood mercury concentration with differences between the males likely being due to the fact that females can expel mercury into the eggs they lay. Juveniles had the lowest blood mercury concentration. Scientists found that the data from juveniles helped to best indicate the local mercury availability as they are fed exclusively from their natal territory.
Elevated levels of mercury have been associated with changes in foraging and brooding behavior among adult common loons, especially in higher concentrations. Studies have found that elevated levels of methylmercury are associated with lethargy and decreased time spent foraging in adult common loons. A different study carried out in the Adirondack Mountains found that elevated levels of methylmercury are associated with reduced diving frequency in adult common loons. One study found that brood productivity was reduced by half when female blood mercury levels exceeded 4.3 μg/g and productivity completely failed when female blood mercury levels exceeded 8.6 μg/g. These results are related to fish mercury levels of 0.21 μg/g and 0.41 μg/g, respectively. As mercury levels and pH are correlated, scientists have found that brood success decreases with decreasing pH such that environments with a pH at around 4.5 exhibited reproductive success below a calculated positive growth rate threshold. An association has also been observed between elevated blood methylmercury levels and aberrant incubation patterns. Adult common loons with high levels of methylmercury were found to spend less time incubating and in the nest, increasing the risks of predation and the eggs overheating/overcooling. Together, the effects of heightened methylmercury levels on parenting behaviors may contribute to lower chick survival rates. One study in Maine and New Hampshire found that high levels of methylmercury in parents are associated with a significant decrease in the number of fledged young, with common loons in the highest risk group producing 41% fewer fledged young than common loons with low levels of methylmercury.
The common loon has also faced a decline in breeding range due to hunting, predation, and water-level fluctuations, or flooding. Some environmentalists attempt to increase nesting success by mitigating the effects of some of these threats, namely terrestrial predation and water-level fluctuations, through the deployment of rafts in the loon's breeding territories. In addition, artificial floating nesting platforms have been provided for the common loon in some lakes to reduce the impact of changing water levels due to dams and other human activities. The common loon abandons lakes that fail to provide suitable nesting habitat due to shoreline development. It is endangered by personal water-craft and powerboats that may drown newly born chicks, wash eggs away, or swamp nests. It is still considered an "injured" species in Alaska as a result of the Exxon Valdez oil spill.
Folk names for the common loon include "big loon", "call-up-a-storm", "greenhead", "hell-diver", "walloon", "black-billed loon", "guinea duck", "imber diver", "ring-necked loon", and "ember-goose".
The common loon was eaten in the Scottish Islands from the Neolithic until the eighteenth century, and its thick layer of fat beneath the skin was used as a cure for sciatica.
Major League Soccer club Minnesota United FC uses the loon in its crest and nickname, as well as a mascot.
Minor League Baseball team Great Lakes Loons, located in Midland, Michigan also uses the loon in its logo and name, as well as a mascot.
Loons are featured prominently in the 1981 film On Golden Pond. Its distinctive sound also appeared in Conan the Barbarian, Out of Africa, Platoon, Guinevere, Harry Potter and the Prisoner of Azkaban, Rick and Morty, Watchmen, Godzilla, Kubo and the Two Strings, , Pet Sematary, 1917 and , among others.
In the 2016 Pixar movie Finding Dory, a somewhat bedraggled and dimwitted loon named Becky is persuaded to use a bucket to help two of the main characters, Nemo and Marlin, get into a marine life institute where the titular Dory is trapped.
Breeding
Habitat selection
Vocalizations
Longevity and terminal investment
Predators and parasites
Status and conservation
Threats to status and conservation
Effects of mercury and acid rain pollution on health and brood productivity
Mercury deposition in the Adirondack Mountains
Breeding range decline
Lead poisoning
In culture
Folklore
Popular culture
Films
Notes
Sources
External links
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